Geographic variation in Crocuta spelaea dentition, beginning from the Middle Pleistocene, can be seen as specialization in western and eastern Eurasia. The sizes of C. spelaea increase from the south to the northwest and northeast. The hyena of the Primorski Krai had the largest teeth.

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Natuurhistorisch Museum Rotterdam

G. Baryshnikov. (1999). Chronological and geographical variability of Crocuta spelaea (Carnivora, Hyaenidae) from the Pleistocene of Russia. Deinsea, 6(1), 155–174.