De geslachten Asperula, Galium, Rubia en Sherardia in Nederland 2. Enige systematische opmerkingen

The opinion of Ehrendorfer & Krendl (1976) concerning the delimitation of the genera Asperula, Cruciata and Galium is accepted by the author. For the Netherlands this means that Asperula odorata, A. glauca and A. humifusa are transferred to Galium. Galium cruciata is transferred to Cruciata. In Galium mollugo no morphological differences could be found between diploid and tetraploid specimens. The chromosome number alone is insufficient to distinguish two species or subspecies in this complex. For the Netherlands three (nomenclaturally unnamed) ecotypes are described. In Galium verum it is possible to recognise two taxa. Both are described as subspecies. One occurring in the dunes is called subsp. maritimum (DC.) Adema stat. Nov., the other occurring in the innermost dunes and also in the rest of the country is called subsp. verum. They key out as follows: Large plants. Stems ascending to almost erect, (15 —)35—85(—120) cm. Leaves (at anthesis) usually much shorter than the internodes. Inflorescens often well developed. ..........subsp. verum Plants much smaller. Stems decumbent-ascending to ascending, sometimes decumbent or creeping, seldom erect, (5—)10—20(—35) cm. Leaves (at anthesis) usually as long as or longer than the internodes. Inflorescens usually small ..................................................... subsp. maritimum In Galium palustre in the Netherlands diploid specimens (subsp. palustre) and octoploid specimens (subsp. elongata) can be found (Kliphuis, 1974). In Sherardia arvensis next to the typical form (var. arvensis) with hairy fruits and deltoidlanceolate, acute calyxteeth, the var. maritima Griseb. with less hairy to almost glabrous fruits and deltoid, obtuse (seldom acute) calyxteeth can be found.