During the germination of barley, the epidermal cells of the scutellum detach themselves from one another and grow to twice their former length. Aleuron proteins are dissolved in the first 3-4 days and subsequently the lipid contents of the spherosomes are digested. This coincides with the development and temporarily increased activity of all metabolic organelles, viz. endoplasmic reticulum, mitochondria, dictyosomes {Golgi apparatus) and leucoplasts. After ten days aqueous vacuoles have developed. After 21 days the cell organelles are smaller and fewer, though some spherosomes are still present.

Acta botanica neerlandica

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Koninklijke Nederlandse Botanische Vereniging

P.J. Nieuwdorp, & Miek C. Buys. (1965). Electron Microscopic Structure of the Epithelial Cells of the Scutellum of Barley II. Cytology of the Cells during Germination. Acta botanica neerlandica, 13(4), 559–565.