The vascular connections between a leaf of the vegetative shoot part and the nectaries of the inflorescence of F. imperialis have been traced with 14C-photosynthates. The conclusion, based on the earlier observed strictly longitudinal translocation of photosynthates in the vascular bundles of the stem, is that the phyllotactical arrangement of the flowers around the stem axis deviates from that of the leaves (2/5) and approximates a 3/8 pattern.

Acta botanica neerlandica

CC BY 3.0 NL ("Naamsvermelding")

Koninklijke Nederlandse Botanische Vereniging

T. Tietema, & J. van Die. (1972). Translocation of assimilates in Fritillaria imperialis L. III. A tracer study on the vascular connections between floral nectaries and leaves. Acta botanica neerlandica, 21(4), 400–403.