The genera Engina and Ameranna nov. gen. (Mollusca: Gastropoda, Buccinoidea, Buccinidae, Pisaniinae) from the Western Atlantic Neogene

The first occurrence of the cosmotropical pisaniine buccinid gastropod genus Engina Gray, 1839 was previously in the lower Gatun Formation (late Miocene) of Panama. Here we describe four new fossil species: E. cantaurana nov. sp., from the early Miocene of Venezuela (the oldest known species of Engina, with an unusually high spire), E. gigas nov. sp., from the late Miocene and early Pliocene of the Dominican Republic (the largest known Engina, with an erect inner lip and obsolete parietal denticle), E. latior nov. sp. from the late Miocene and early Pliocene of the Dominican Republic (a more typical species of Engina in terms of apertural dentition) and Engina moinensis nov. sp from the Pleistocene of Costa Rica and the Bocas del Toro Area (a species with possible paciphile affinities). We propose the new pisaniine genus Ameranna (type species: Anna florida García, 2008) for four living Caribbean species and for two fossil species: A. primitiva nov. sp., from the early Miocene of Venezuela (the oldest known species of Ameranna) and the new Pliocene species A. minuscula nov. sp. from the Dominican Republic. The new genus differs from the early Oligocene to Recent eastern Atlantic genus Anna by having a lirate rather than denticulate outer lip.