Patterns of distribution and territoriality in the two male forms of Mnais pruinosa pruinosa Selys (Zygoptera: Calopterygidae)

In the study area 2 forms of M. p. pruinosa males (f. esakii and f. strigata) occurred. Immature damselflies dispersed away from the stream. After maturation they returned to the stream and often moved along the stream searching for territory sites. Once esakii males established their territories, strigata males stettled around these territories as satellites. The males of esakii had many mating chances, utilizing their territories. Male movements and settlement were influenced by fighting among the males and also by the frequency of female appearances. Both esakii and strigata males aggregated in the habitat sections where many females appeared. Mating strategies of esakii and strigata males are described as territorial-guarding and satellite-multicopulation (nonguarding), respectively. A possible mechanism for coexistence of both types of males is suggested.