Notes on Buliminidae, 5. On the systematic position of Arabian Buliminidae (Gastropoda Pulmonata), with the description of a new genus

It is argued that the Chondrulinae are of a polyphyletic origin: the penial appendix of the Buliminidae has been lost independently in at least six different lineages. The Azorean species, which in contrast to the Canarian Buliminidae lack a penial appendix, are assigned to Macaronapaeus. Spelaeoconcha is removed from the Buliminidae; its systematic affinities within the Pupilloidea remains however unclear. A new division of the Buliminidae into two subfamilies is given: Bulimininae and Eninae. The former is mainly characterised by the presence of a penial caecum and contains the genera Buliminus, Pene, Paramastus (with the subgenera Paramastus s. str. and Cyrenaeus) and the here described Mordania (with the monotypic subgenera Mordania s. str. and Iranopsis). The systematic position of the four Arabian Buliminidae is discussed. The two south-western species hedjazicus and sabaeanus belong to Paramastus whereas the two south-eastern taxa omanensis and jousseaumei are classified within Mordania and Pseudonapaeus, respectively. The last two species are of turanian origin.